- Despite the small sample set from Neolithic Bulgaria and western Ukraine, two out of the four haplogroups are shared with Steppe_EMBA, including the fairly specific H2a2 - Steppe_EMBA potentially shares very deep ancestry with Iran_Neolithic, via basal clades such as T2c and possibly X2; these markers could have arrived on the steppe in any number of ways, and indeed may have been sitting there since the Mesolithic - But Iran_Chalcolithic is still a horrible match with Steppe_EMBA, and Maikop no betterNeedless to say, I'm looking forward to those new Neolithic Balkan samples from the upcoming Mathieson et al. paper (see here). I'm pretty sure now that at least some Steppe_EMBA groups will show admixture from Neolithic or Chalcolithic populations from the eastern Balkans. See also... Mixed marriages on the early Eneolithic steppe Modeling Steppe_EMBA Male-dominated conquest of Europe by Bronze Age steppe pastoralists The story of mtDNA haplogroup U7
Friday, October 28, 2016
I updated my table of Steppe_EMBA mitochondrial (mtDNA) haplogroups with data from Wilde 2014, Broushaki et al. 2016, Sokolov et al. 2016, Jones et al. 2015, and extra data from Lazaridis et al. 2016. The accompanying map is from Wilde 2014 and shows sampling locations of many of the individuals in the spreadsheet. here). At least now I can see a few shared haplogroups. Some thoughts:
Thursday, October 27, 2016
Below is another abstract from the upcoming 6th DNA Polymorphisms in Human Populations conference in Paris. It was added to the abstract book after I first blogged about the conference here. The authors are probably first testing mtDNA to check which of the samples have enough DNA for full genome sequencing or enrichment capture of genome-wide SNPs. So the fact that they managed to sequence so many mitogenomes means that we might soon see genome-wide data for most of the same samples. The results make sense considering the genetic structure of present-day Armenians (see here). Indeed, I suspect that early Neolithic farmers from the southern Caucasus will come out looking very similar overall to present-day Armenians. If so, this will probably be hailed by many as evidence supporting the Armenian Plateau Indo-European homeland theory. However, as I've already shown, it's very likely that the Armenian Plateau was affected by population movements from the Eastern European steppe during the Bronze Age which may have introduced Indo-European languages to the region (see here).
The origin of the Armenian people is heavily debated among historians and archaeologists. Despite a long history and vast archaeological records in Armenia, it has proven very challenging to infer the demographic events that led to the formation of Armenians as a distinct ethno-cultural group. To obtain a detailed understanding of the demographic events in Armenia across millennia, we study complete mitochondrial genomes from 49 ancient individuals covering 7800 years and compare them with that of modern Armenians (n=206) and seven neighboring populations (n=482). In this context, the lowest genetic distance was observed between the modern and ancient Armenians and this was also reflected in network analyses and discriminant analysis of principal components (dapc) showing genetic proximity between the ancient individuals and modern Armenians. We used Approximate Bayesian Computation (ABC) to test five different demographic scenarios of the Armenian population, and the simulations favored a model where both ancient and modern Armenians derive from the same source population. We conclude that there is a strong signal of continuity in the maternal Armenian gene pool during the last 7800 years.Margaryan et al., 7800 years of Mitochondrial genetic continuity in Armenia, 6th DNA Polymorphisms in Human Populations, Talk Workhsop Genomic Demography, Musee de l’Homme, Paris, 7-10 December, 2016
Thursday, October 20, 2016
From The Siberian Times:
He has a massive nose with flared nostrils, wide open mouth, a bushy moustache and a beard. And yet all is not quite as it seems, for this sculpture, the most northerly of this genre in Asia, underwent an historic version of plastic surgery perhaps 1,500 years ago to give him a less Caucasian and more Asian appearance, according to experts. ... So the original European look of the idol was changed to a more Asian countenance. Why would this happen? 'Judging by archeological finds found inside the grottos, this anthropomorphic idol was made during the Scythian time,' Yuri Grevtsov said. 'The first change came when the more European looking face was transformed to make it appear more Mongoloid was likely to have happened in the early Middle Ages with a shift of the population in the Angara River area,' he said. In other words, incoming ethnic groups preferred the idol to be more akin to their own looks.Source: Siberia's stone idols - 2,400 year old Ust-Taseyevsky idol 'underwent racial realignment early in Middle Ages', losing his European looks by Tamara Zubchuk
Friday, October 14, 2016
For your pleasure and my satisfaction: a nice little slide show on the ancient population history of South Asia. Click on the first image to get started. The images are based on my latest Principal Component Analysis (PCA) of the world (see here). Any other questions? Ask in the comments. Caste is in the genes
Below is my new Principal Component Analysis (PCA) or genetic map of global human population structure. I think it's a little bit special, and we can discuss why in the comments if anyone's interested. The datasheet is available here; it can be used to generate 2D and 3D PCA plots, and to model samples of your choice using the nMonte and 4mix R scripts.
Karitiana Ulchi 64.45 AfontovaGora3 34.2 Dai 1.35 distance%=0.5479/distance=0.005479 Wichi Ulchi 66.8 AfontovaGora3 33.2 Dai 0 distance%=0.5055/distance=0.005055 Kalash Iran_Neolithic:I1945 38.6 Paniya 20.2 Yamnaya_Samara:I0357 17.4 Afanasievo:RISE509 16.55 Andronovo:RISE505 3.8 Iran_Late_Neolithic:I1671 3.45 Iran_Hotu:I1293 0 distance%=0.3793/distance=0.003793 Brahmin_Uttar_Pradesh Paniya 54 Yamnaya_Samara:I0357 24.55 Iran_Neolithic:I1945 21.45 Afanasievo:RISE509 0 Andronovo:RISE505 0 Iran_Hotu:I1293 0 Iran_Late_Neolithic:I1671 0 distance%=0.997/distance=0.00997 Brahmin_Tamil_Nadu Paniya 57.2 Iran_Neolithic:I1945 27.7 Yamnaya_Samara:I0357 15.1 Afanasievo:RISE509 0 Andronovo:RISE505 0 Iran_Hotu:I1293 0 Iran_Late_Neolithic:I1671 0 distance%=1.0312/distance=0.010312 Pathan Paniya 24.9 Iran_Neolithic:I1945 23.15 Andronovo:RISE505 19.95 Iran_Late_Neolithic:I1671 17.85 Yamnaya_Samara:I0357 14.15 Afanasievo:RISE509 0 Iran_Hotu:I1293 0 distance%=0.5729/distance=0.005729It's interesting to note that the Dai from southern China help to improve the fit for Karitiana from the Amazon basin, but not the Wichi from Argentina. Also, Andronovo significantly improves the fit for the East Iranian Pathans or Pashtuns, but clearly not as much for the Indo-Aryan Kalash, and none at all for Brahmins from India, who are also Indo-Aryans. Why? Don't know, but it might well be an important question in regards to the origins and spread of Indo-Iranian languages. See also... The peopling of South Asia: an illustrated guide
Monday, October 10, 2016
Iron Age European-like people in what is now western China buried their dead with cannabis plants. Courtesy of Phys.org:
The skeleton has been identified as once belonging to a Caucasian man approximately 35 years old at the time of his death. Those that had buried him had placed a willow pillow under his head and had then placed a shroud of (13) cannabis plants over his chest reaching from below his pelvis at one end to the side of his face on the other. The skeleton lay in one of the 240 graves in the area known as the Jiayi cemetery. The people that lived in the area at the time were part of a Kingdom from 3,000 and 2,000 years ago known as the Subeixi. Prior research has shown the people lived there because it was an oasis in the desert, one that had become an important place for travelers to rest during their trek along the Silk Road.Full article: Ancient skeleton covered in cannabis shroud unearthed in China Based on this paper at Economic Botany:
Abstract: An extraordinary cache of ancient, well-preserved Cannabis plant remains was recently discovered in a tomb in the Jiayi cemetery of Turpan, NW China. Radiometric dating of this tomb and the archeobotanical remains it contained indicate that they are approximately 2800–2400 years old. Both morphological and anatomical features support the identification of the plant remains as Cannabis. Research discussed in this paper describes 13 nearly whole plants of Cannabis that appear to have been locally produced and purposefully arranged and used as a burial shroud which was placed upon a male corpse. This unique discovery provides new insight into the ritualistic use of Cannabis in prehistoric Central Eurasia. Furthermore, the fragmented infructescences of Cannabis discovered in other tombs of the Jiayi cemetery, together with similar Cannabis remains recovered from coeval tombs in the ancient Turpan cemetery along with those found in the Altai Mountains region, reveal that Cannabis was used by the local Central Eurasian people for ritual and/or medicinal purposes in the first millennium before the Christian era.Hongen Jiang et al., Ancient Cannabis Burial Shroud in a Central Eurasian Cemetery, Economic Botany (2016). DOI: 10.1007/s12231-016-9351-1 See also... Bronze Age dope dealers
Saturday, October 8, 2016
Max Planck's Johannes Krause featured this curious map at a recent talk in Moscow on the Proto-Indo-European homeland debate (two hours into the clip here). The story of mtDNA haplogroup U7
Tuesday, October 4, 2016
Here's another graph based on my new D-stats datasheet. The contrast in the population affinities of Armenia_MLBA (Middle Late Bronze Age) and Armenia_EBA (Early Bronze Age) is, at least for me, surprising.
Yoruba Sintashta Armenia_EBA Armenia_MLBA f4 0.000676 Z 1.395 Yoruba Potapovka Armenia_EBA Armenia_MLBA f4 0.000459 Z 1.275 Yoruba Corded_Ware_CE Armenia_EBA Armenia_MLBA f4 0.000383 Z 0.933 Yoruba Poltavka Armenia_EBA Armenia_MLBA f4 0.000373 Z 0.814 Yoruba Andronovo Armenia_EBA Armenia_MLBA f4 0.000214 Z 0.486By the way, these stats are based on transversion sites only to limit the effects of post-mortem damage on the ancient samples, some of which are not UDG treated. Update 06/10/2016: As far as I can see, the qpAdm modeling software shows that Sintashta is indeed the best available proxy for the European-like admixture in Armenia_MLBA.
Outgroups Andamanese_Onge Bichon Chukchi Han Israel_Natufian Karitiana Kostenki14 MA1 Mbuti Papuan Ust_Ishim Armenia_MLBA Armenia_EBA 0.799±0.069 Sintashta 0.201±0.069 chisq 7.181 tail prob 0.618257 Armenia_MLBA Armenia_EBA 0.835±0.068 Andronovo (3) 0.165±0.068 chisq 9.549 tail prob 0.388179 Armenia_MLBA Armenia_EBA 0.842±0.065 Andronovo (4) 0.158±0.065 chisq 9.742 tail prob 0.371809 Armenia_MLBA Armenia_EBA 0.838±0.069 Srubnaya 0.162±0.069 chisq 9.993 tail prob 0.351059Update 08/10/2016: By the way, the fact that Kura-Araxes shares high genetic drift with many Indo-European-speaking Southern Europeans, such as Albanians, has no bearing on its posited identity as an Hurrian-speaking population. That's because this inflated genetic affinity is mediated via ancient groups of largely Near Eastern origin not directly related to Kura-Araxes, such as Copper and Bronze Age pre-Indo-European Europeans. On the other hand, Caucasians, particularly Northeast Caucasians, in all likelihood do share direct ancestry with Kura-Araxes.
Monday, October 3, 2016
I've put together a new D-stats sheet that might be useful in the Indo-European homeland debate (see here). It features new samples from the EGDP dataset, with most of the stats based on over 750K SNPs. The stats are of the form D(Chimp,Ancient)(Mbuti,X). The idea that Indo-Iranian languages arrived in Central and South Asia from the Armenian Plateau and/or eastern Anatolia during the Bronze Age, as per Gamkrelidze and Ivanov (check out their article here and, if you don't have access, crazy map here), is still popular with a lot of people. But it's most certainly a dud. There's too much Bronze Age steppe ancestry in this part of the world, particularly among the more isolated Indo-Iranian populations like Pamir Tajiks and the Kalasha, as well as upper caste Indians, to ignore. At the same time, there is no hard data linking any of these groups to Bronze Age Armenia or Anatolia. here), which they clearly should if their Indo-European ancestors migrated en masse from Transcaucasia to the Pontic-Caspian steppe in an anti-clockwise direction around the Caspian Sea. here). And, judging by the affinities of CHG and ancient groups in large part of CHG origin, these people were more likely the speakers of Caucasian languages than of Proto-Indo-European. Hurrians and the others